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Post by clouddesignc7 on Sept 30, 2016 15:37:07 GMT -5
On mtDNA M (and so on M1 aswell):
The eastern-African group, named M1, is characterized within M by a consensus HVS-I motif defined by four transitions at nt 16,129, 16,189, 16,249 and 16,311. This HVS-I signature motif is not found either in our or other Indian samples - Semino et al
We found 489C (Table 3) in all Indian and eastern-African haplogroup M mtDNAs analysed, but not in the non-M haplogroup controls, including 20 Africans representing all African main lineages (6 L1, 4 L2, 10 L3) and 11 Asians. These findings, and the lack of positive evidence (given the RFLP status) that the 10400 CT transition defining M has happened more than once, suggest that it has a single common origin, but do not resolve its geographic origin. Analysis of position 10873 (the MnlI RFLP) revealed that all the M molecules (eastern African, Asian and those sporadically found in our population surveys) were 10873C (Table 3). As for the non-M mtDNAs, the ancient L1 and the L2 African-specific lineages5, as well as most L3 African mtDNAs, also carry 10873C.
Conversely,non-M mtDNAs, the ancient L1 and the L2 African-specific lineages5, as well as most L3 African mtDNAs, also carry 10873C. Conversely, all non-M mtDNAs of non-African origin analysed so far carry 10873T. These data indicate that the **transition 10400 C-->T, which defines haplogroup M, arose on an African background characterized by the ancestral state 10873C**, which is also present in four primate (common and pygmy chimps, gorilla and orangutan) mtDNA sequences. - Semino et al
M arose 60,000 years ago.
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Post by clouddesignc7 on Oct 6, 2016 9:21:00 GMT -5
haplogroup M originated approximately 60,000 years ago
- Murci et al 1999
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Post by clouddesignc7 on Oct 6, 2016 9:56:26 GMT -5
In regards to mtDNA: " Egypt’s location and amalgam of past ruling populations mixing with the Egyptians has resulted in a heterogeneous make-up. The haplotype breakdown of this dataset is predominantly European (67.5%), followed by African (20.6%) and Asian (11.9%). The breakdown is as follows: European origin (n = 187) including R0 and its subgroups (31.4%), I (3.2%), J (7.6%), K (4.7%), T (9.4%), U (9.0%), W (0.7%), X (1.4%); African origin (n = 57) including L0 (2.2%), L1 (2.5%), L2 (3.6%), L3 (12.3%); and Asian origin (n = 33) including M (6.9%), N (5.1%)."--Source: Mitochondrial control region sequences from an Egyptian population sample Jessica L. Saunier et al. 2009 Download link: www.sendspace.com/file/fk714r Sample consisted of males from Alexandria. So, the heterogeneity can be attributed to the invasions throughout Egyptian history. Aside perhaps for M1 which has an possible/probable African origin
So now it's probably European (58.5%), African (36.5%) and Asian (5.1%).
Since M is African and since the U might be U6, and that's African.
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Post by clouddesignc7 on Oct 6, 2016 12:47:15 GMT -5
Based on the high frequency and diversity of haplogroup M in India and elsewhere in Asia, some authors have suggested (versus [3]) that M may have arisen in Southwest Asia [16,17,31]. Finding M1 or a lineage ancestral to M1 in India, could help to explain the presence of M1 in Africa as a result of a back migration from India. Yet, to date this has not been achieved [15], this study). Therefore, one cannot rule out the still most parsimonious scenario that haplogroup M arose in East Africa [3]. Furthermore, the lack of L3 lineages other than M and N (indeed, L3M and L3N) in India is more consistent with the African launch of haplogroup M. On the other hand, one also observes that: i) M1 is the only variant of haplogroup M found in Africa; ii) M1 has a fairly restricted phylogeography in Africa, barely penetrating into sub-Saharan populations, being found predominantly in association with the Afro-Asiatic linguistic phylum – a finding that appears to be inconsistent with the distribution of sub-clades of haplogroups L3 and L2 that have similar time depths. That, plus the presence of M1 without accompanying L lineages in the Caucasus [32] and [our unpublished data], leaves the question about the origin of haplogroup M still open.
- Metspalu et al 2004, Most of the extant mtDNA boundaries in South and Southwest Asia were likely shaped during the initial settlement of Eurasia by anatomically modern humans
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Post by clouddesignc7 on Dec 27, 2016 9:48:04 GMT -5
On E-M2 E1b1a:
"The analysis of our data provides further evidence for the homogeneity of the Y chromosome gene pool of sub-Saharan West Africans, due to the high frequency of haplogroup E3a-M2. Its frequency and diversity in West Africa are among the highest found, suggesting an early local origin and expansion in the last 20–30 ky.
[...]
The E3a*-M2 microsatellite profiles of Mandenka and Balanta are the most diverse among our data (RST average gene diversity, see Additional file 2) and attest to an earlier origin or more pronounced expansion. Since the corresponding parameter in Fulbe is less diverse we consider this to signal either a genetic bottleneck or their more recent expansion and late arrival in the West. The data are consistent with the less diverse E3a-M2 profile in Central and South Africans (data not shown).
[...]
The predominance and high diversity of haplogroup E3a*-M2 suggests a demographic expansion in the equatorial western fringe [of West Africa / Guinea Bissau], possibly supported by a local agricultural center. The paternal pool of the Mandenka and Balanta displays evidence of a particularly marked population growth among the Guineans, possibly reflecting the demographic effects of the agriculturalist lifestyle and their putative relationship to the people that introduced early cultivation practices into West Africa."
- Rosa et al 2007, Y-chromosomal diversity in the population of Guinea-Bissau: a multiethnic perspective
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Post by clouddesignc7 on Dec 27, 2016 10:41:38 GMT -5
On the E-M35 -carrying Neolithic Middle Easterners (the Natufians -- Neolithic means farmers) these studies keep mentioning, who went into Europe:
"If the late Pleistocene Natufian sample from Israel is the source from which that Neolithic spread was derived, there was clearly a sub-Saharan African element present of almost equal importance as the Late Prehistoric Eurasian element."
- Brace et al 2005, The Questionable contribution of Neolithic to Bronze age European craniofacial form
Whole thing:
"The surprise is that the Neolithic peoples of Europe and their Bronze Age successors are not closely related to the modern inhabitants although the prehistoric/modern ties are somewhat more apparent in southern Europe. It is a further surprise that the Epipalaeolithic Natufian of Israel from whom the Neolithic realm was assumed to arise has a clear link to sub-Saharan Africa."
[...]
"The assessment of prehistoric and recent human craniofacial dimensions supports the picture documented by genetics that the extension of Neolithic agriculture from the Near East westward to Europe and across North Africa was accomplished by a process of demic diffusion (11–15). If the Late Pleistocene Natufian sample from Israel is the source from which that Neolithic spread was derived, then there was clearly a SubSaharan African element present of almost equal importance as the Late Prehistoric Eurasian element. At the same time, the failure of the Neolithic and Bronze Age samples in central and northern Europe to tie to the modern inhabitants supports the suggestion that, while a farming mode of subsistence was spread westward and also north to Crimea and east to Mongolia by actual movement of communities of farmers, the indigenous foragers in each of those areas ultimately absorbed both the agricultural subsistence strategy and also the people who had brought it. The interbreeding of the incoming Neolithic people with the in situ foragers diluted the Sub-Saharan traces that may have come with the Neolithic spread so that no discoverable element of that remained. This picture of a mixture between the incoming farmers and the in situ foragers had originally been supported by the archaeological record alone (6, 9, 33, 34, 48, 49), but this view is now reinforced by the analysis of the skeletal morphology of the people of those areas where prehistoric and recent remains can be metrically compared."
- Brace et al 2005
"One can identify Negroid traits of nose and prognathism appearing in Natufian latest hunters (McCown, 1939) and in Anatolian and Macedonian first farmers (Angel, 1972), probably from Nubia via the predecesors of the Badarians and Tasians"
- Larry Angel
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Post by clouddesignc7 on Dec 27, 2016 11:09:28 GMT -5
More on the Natufians (the first farmers), Southeast Europe, and sub-Saharan Africa:
F. X. Ricaut M. Waelkens Cranial Discrete Traits in a Byzantine Population and Eastern Mediterranean Population Movements Human Biology - Volume 80, Number 5, October 2008, pp. 535-564
Abstract: "Since the beginning of the Holocene, the Anatolian region has been a crossroads for populations and civilizations from Europe, Asia, and the Near to Middle East, with increasing interactions since the Bronze Age. In this context, we examine cranial discrete traits from a Byzantine population from southwest Turkey, excavated at the archeological site of Sagalassos; the site displays human occupation since the 12th millennium b.p. To investigate the biological history of this population, we analyzed the frequency distribution of 17 cranial discrete traits from Sagalassos and 27 Eurasian and African populations. Ward’s clustering procedure and multidimensional scaling analyses of the standardized mean measure of divergence (MMDst), based on trait frequencies, were used to represent the biological affinity between populations. Our results, considered within a large interpretive framework that takes into account the idea that populations are dynamic entities affected by various influences through time and space, revealed different strata of the Sagalassos biological history. Indeed, beyond an expected biological affinity of the Sagalassos population with eastern Mediterranean populations, we also detected affinities with sub-Saharan and northern and central European populations. We hypothesize that these affinity patterns in the Sagalassos biological package are the traces of the major migratory events that affected southwest Anatolia over the last millennia, as suggested from biological, archeological, and historical data."
"Keeping in mind these three elements, if we consider the affinity of the Sagalassos population with the sub-Saharan populations from Gabon and Somalia, a recent direct contact between these populations and regions probably can be excluded because they are seperated by significant geographic distances. However, indirect contacts through geographically intermediary populations carrying "sub-Saharan" biological features in the late Pleistocene-Holocene period are discussion points."
"From the Mesolithic to the early Neolithic period different lines of evidence support an out-of-Africa Mesolithic migration to the Levant by northeastern African groups that had biological affinities with sub-Saharan populations. From a genetic point of view, several recent genetic studies have shown that sub-Saharan genetic lineages (affiliated with the Y-chromosome PN2 clade; Underhill et al. 2004) have spread through Egypt into the Near East, the Mediterranean area, and, for some lineages, as far north as Turkey(E3b-M35 Y lineage; Cinnioglu et al. 2004; Luis et al. 2004), probably during several dispersal episodes since the Mesolithic (Cinnioglu et al. 2004; King et al. 2008; Lucotte and Mercier 2003; Luis et al. 2004; Quintanna-Murci et al. 1999; Semino et al. 2004; Underhill et al. 2001). This finding is in agreement with morphological data that suggest that populations with sub-Saharan morphological elements were present in northeastern Africa, from the Paleolithic to at least the early Holocene, and diffused northward to the Levant and Anatolia beginning in the Mesolithic. Indeed, the rare and incomplete 33,000-year-old Nazlet Khater specimen (Pinhasi and Semal 2000), the Wadi Kubbaniya skeleton from the late Paleolithic site in the upper Nile Valley (Wendorf et al. 1986), the Qarunian (Faiyum) early Neolithic crania (Henneberg et al. 1989; Midant-Reynes 2000), and the Nabta specimen from the Neolithic Nabta Playa site in the western desert of Egypt (Henneberg et al. 1980)-show, with regard to the great African biological diversity, similarities with some of the sub-Saharan middle Paleolithic and modern sub-Saharan specimens. This affinity pattern between ancient Egyptians and sub-Saharans has also been noticed by several other investigators (Angel 1972; Berry and Berry 1967, 1972; Keita 1995) and has been recently reinforced by the study of Brace et al. (2005), which clearly shows that the cranial morphology of prehistoric and recent northeast African populations is linked to sub-Saharan populations (Niger-Congo populations). These results support the hypothesis that some of the Paleolithic-early Holocene populations from northeast Africa were probably descendents of sub-Saharan ancestral populations."
"A late Pleistocene-early Holocene northward migration (from Africa to the Levant and to Anatolia) of these populations has been hypothesized from skeletal data (Angel 1972, 1973; Brace 2005) and from archaeological data, as indicated by the probable Nile Valley origin of the "Mesolithic" (epi-Paleolithic) Mushabi culture found in the Levant (Bar Yosef 1987). This migration finds some support in the presence in Mediterranean populations (Sicily, Greece, southern Turkey, etc.; Patrinos et al.; Schiliro et al. 1990) of the Benin sickle cell haplotype. This haplotype originated in West Africa and is probably associated with the spread of malaria to southern Europe through an eastern Mediterranean route (Salares et al. 2004)following the expansion of both human and mosquito populations brought about by the advent of the Neolithic transition (Hume et al 2003; Joy et al. 2003; Rich et al 1998). This northward migration of northeastern African populations carrying sub-Saharan biological elements is concordant with the morphological homogeneity of the Natufian populations (Bocquentin 2003), which present morphological affinity with sub-Saharan populations (Angel 1972; Brace et al. 2005). In addition, the Neolithic revolution was assumed to arise in the late Pleistocene Natufians and subsequently spread into Anatolia and Europe (Bar-Yosef 2002), and the first Anatolian farmers, Neolithic to Bronze Age Mediterraneans and to some degree other Neolithic-Bronze Age Europeans, show morphological affinities with the Natufians (and indirectly with sub-Saharan populations; Angel 1972; Brace et al 2005), in concordance with a process of demic diffusion accompanying the extension of the Neolithic revolution (Cavalli-Sforza et al. 1994)."
"Following the numerous interactions among eastern Mediterranean and Levantine populations and regions, caused by the introduction of agriculture from the Levant into Anatolia and southeastern Europe, there was, beginning in the Bronze Age, a period of increasing interactions in the eastern Mediterranean, mainly during the Greek, Roman, and Islamic periods. These interactions resulted in the development of trading networks, military campaigns, and settler colonization. Major changes took place during this period, which may have accentuated or diluted the sub-Saharan components of earlier Anatolian populations. The second option seems more likely, because even though the population from Sagalassos territory was interacting with northeastern African and Levantine populations [trade relationships with Egypt (Arndt et al. 2003), involvement of thousands of mercanries from Pisidia (Sagalassos region) in the war around 300 B.C. between the Ptolemaic kingdom (centered in Egypt) and the Seleucid kingdom (Syria/Mesopotamia/Anatolia), etc.], the major cultural and population interactions involving the Anatolian populations since the Bronze Age occured with the Mediterranean populations form southeastern Europe, as suggested from historical and genetic data."
"Consequently, one may hypothesize as the most parsimonious explanation that sub-Saharan biological elements were introduced into the Anatolian populations after the Neolithic spread and have been preserved since this time, at least until the 11th-13th century A.D., in the population living in the Sagalassos territory of southwestern Anatolia. This scenario implies that the affinity between Sagalassos and the two sub-Saharan populations (Gabon and Somalia) is more likely due to the sharing of a common ancestor and that the major changes and increasing interactions in the eastern Mediterranean beginning in the Bronze Age did not erase some of the sub-Saharan elements carried by Anatolian populations, as shown by genetic data and the morphologivcal features of our southwestern Anatolian sample."
"In this context it is likely that Bronze Age events may have facilitated the southward diffusion of populations carrying northern and central European biological elements and may have contributed to some degree of admixture between northern and central Europeans and Anatolians, and on a larger scale, between northeastern Mediterraneans and Anatolians. Even if we do not know which populations were involved, historical and archaeological data suggest, for instance, the 2nd millenium B.C. Minoan and later Mycenaean occupation of Anatolian coast, the arrival in Anatolia in the early 1st millennium B.C. of the Phrygians coming from Thrace, and later the arrival of settlers from Macedonia in Pisidia and in the Sagalassos territory (under Seleucid rule). The coming of the Dorians from Northern Greece and central Europe (the Dorians are claimed to be one of the main groups at the origin of the ancient Greeks) may have also brought northern and central European biological elements into southern populations. Indeed, the Dorians may have migrated southward to the Peloponnese, across the southern Aegean and Create, and later reached Asia Minor."
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Post by history91 on Dec 30, 2016 15:54:37 GMT -5
More on the Natufians (the first farmers), Southeast Europe, and sub-Saharan Africa: F. X. Ricaut M. Waelkens Cranial Discrete Traits in a Byzantine Population and Eastern Mediterranean Population Movements Human Biology - Volume 80, Number 5, October 2008, pp. 535-564 Abstract: "Since the beginning of the Holocene, the Anatolian region has been a crossroads for populations and civilizations from Europe, Asia, and the Near to Middle East, with increasing interactions since the Bronze Age. In this context, we examine cranial discrete traits from a Byzantine population from southwest Turkey, excavated at the archeological site of Sagalassos; the site displays human occupation since the 12th millennium b.p. To investigate the biological history of this population, we analyzed the frequency distribution of 17 cranial discrete traits from Sagalassos and 27 Eurasian and African populations. Ward’s clustering procedure and multidimensional scaling analyses of the standardized mean measure of divergence (MMDst), based on trait frequencies, were used to represent the biological affinity between populations. Our results, considered within a large interpretive framework that takes into account the idea that populations are dynamic entities affected by various influences through time and space, revealed different strata of the Sagalassos biological history. Indeed, beyond an expected biological affinity of the Sagalassos population with eastern Mediterranean populations, we also detected affinities with sub-Saharan and northern and central European populations. We hypothesize that these affinity patterns in the Sagalassos biological package are the traces of the major migratory events that affected southwest Anatolia over the last millennia, as suggested from biological, archeological, and historical data."
"Keeping in mind these three elements, if we consider the affinity of the Sagalassos population with the sub-Saharan populations from Gabon and Somalia, a recent direct contact between these populations and regions probably can be excluded because they are seperated by significant geographic distances. However, indirect contacts through geographically intermediary populations carrying "sub-Saharan" biological features in the late Pleistocene-Holocene period are discussion points."
"From the Mesolithic to the early Neolithic period different lines of evidence support an out-of-Africa Mesolithic migration to the Levant by northeastern African groups that had biological affinities with sub-Saharan populations. From a genetic point of view, several recent genetic studies have shown that sub-Saharan genetic lineages (affiliated with the Y-chromosome PN2 clade; Underhill et al. 2004) have spread through Egypt into the Near East, the Mediterranean area, and, for some lineages, as far north as Turkey(E3b-M35 Y lineage; Cinnioglu et al. 2004; Luis et al. 2004), probably during several dispersal episodes since the Mesolithic (Cinnioglu et al. 2004; King et al. 2008; Lucotte and Mercier 2003; Luis et al. 2004; Quintanna-Murci et al. 1999; Semino et al. 2004; Underhill et al. 2001). This finding is in agreement with morphological data that suggest that populations with sub-Saharan morphological elements were present in northeastern Africa, from the Paleolithic to at least the early Holocene, and diffused northward to the Levant and Anatolia beginning in the Mesolithic. Indeed, the rare and incomplete 33,000-year-old Nazlet Khater specimen (Pinhasi and Semal 2000), the Wadi Kubbaniya skeleton from the late Paleolithic site in the upper Nile Valley (Wendorf et al. 1986), the Qarunian (Faiyum) early Neolithic crania (Henneberg et al. 1989; Midant-Reynes 2000), and the Nabta specimen from the Neolithic Nabta Playa site in the western desert of Egypt (Henneberg et al. 1980)-show, with regard to the great African biological diversity, similarities with some of the sub-Saharan middle Paleolithic and modern sub-Saharan specimens. This affinity pattern between ancient Egyptians and sub-Saharans has also been noticed by several other investigators (Angel 1972; Berry and Berry 1967, 1972; Keita 1995) and has been recently reinforced by the study of Brace et al. (2005), which clearly shows that the cranial morphology of prehistoric and recent northeast African populations is linked to sub-Saharan populations (Niger-Congo populations). These results support the hypothesis that some of the Paleolithic-early Holocene populations from northeast Africa were probably descendents of sub-Saharan ancestral populations."
"A late Pleistocene-early Holocene northward migration (from Africa to the Levant and to Anatolia) of these populations has been hypothesized from skeletal data (Angel 1972, 1973; Brace 2005) and from archaeological data, as indicated by the probable Nile Valley origin of the "Mesolithic" (epi-Paleolithic) Mushabi culture found in the Levant (Bar Yosef 1987). This migration finds some support in the presence in Mediterranean populations (Sicily, Greece, southern Turkey, etc.; Patrinos et al.; Schiliro et al. 1990) of the Benin sickle cell haplotype. This haplotype originated in West Africa and is probably associated with the spread of malaria to southern Europe through an eastern Mediterranean route (Salares et al. 2004)following the expansion of both human and mosquito populations brought about by the advent of the Neolithic transition (Hume et al 2003; Joy et al. 2003; Rich et al 1998). This northward migration of northeastern African populations carrying sub-Saharan biological elements is concordant with the morphological homogeneity of the Natufian populations (Bocquentin 2003), which present morphological affinity with sub-Saharan populations (Angel 1972; Brace et al. 2005). In addition, the Neolithic revolution was assumed to arise in the late Pleistocene Natufians and subsequently spread into Anatolia and Europe (Bar-Yosef 2002), and the first Anatolian farmers, Neolithic to Bronze Age Mediterraneans and to some degree other Neolithic-Bronze Age Europeans, show morphological affinities with the Natufians (and indirectly with sub-Saharan populations; Angel 1972; Brace et al 2005), in concordance with a process of demic diffusion accompanying the extension of the Neolithic revolution (Cavalli-Sforza et al. 1994)."
"Following the numerous interactions among eastern Mediterranean and Levantine populations and regions, caused by the introduction of agriculture from the Levant into Anatolia and southeastern Europe, there was, beginning in the Bronze Age, a period of increasing interactions in the eastern Mediterranean, mainly during the Greek, Roman, and Islamic periods. These interactions resulted in the development of trading networks, military campaigns, and settler colonization. Major changes took place during this period, which may have accentuated or diluted the sub-Saharan components of earlier Anatolian populations. The second option seems more likely, because even though the population from Sagalassos territory was interacting with northeastern African and Levantine populations [trade relationships with Egypt (Arndt et al. 2003), involvement of thousands of mercanries from Pisidia (Sagalassos region) in the war around 300 B.C. between the Ptolemaic kingdom (centered in Egypt) and the Seleucid kingdom (Syria/Mesopotamia/Anatolia), etc.], the major cultural and population interactions involving the Anatolian populations since the Bronze Age occured with the Mediterranean populations form southeastern Europe, as suggested from historical and genetic data."
"Consequently, one may hypothesize as the most parsimonious explanation that sub-Saharan biological elements were introduced into the Anatolian populations after the Neolithic spread and have been preserved since this time, at least until the 11th-13th century A.D., in the population living in the Sagalassos territory of southwestern Anatolia. This scenario implies that the affinity between Sagalassos and the two sub-Saharan populations (Gabon and Somalia) is more likely due to the sharing of a common ancestor and that the major changes and increasing interactions in the eastern Mediterranean beginning in the Bronze Age did not erase some of the sub-Saharan elements carried by Anatolian populations, as shown by genetic data and the morphologivcal features of our southwestern Anatolian sample."
"In this context it is likely that Bronze Age events may have facilitated the southward diffusion of populations carrying northern and central European biological elements and may have contributed to some degree of admixture between northern and central Europeans and Anatolians, and on a larger scale, between northeastern Mediterraneans and Anatolians. Even if we do not know which populations were involved, historical and archaeological data suggest, for instance, the 2nd millenium B.C. Minoan and later Mycenaean occupation of Anatolian coast, the arrival in Anatolia in the early 1st millennium B.C. of the Phrygians coming from Thrace, and later the arrival of settlers from Macedonia in Pisidia and in the Sagalassos territory (under Seleucid rule). The coming of the Dorians from Northern Greece and central Europe (the Dorians are claimed to be one of the main groups at the origin of the ancient Greeks) may have also brought northern and central European biological elements into southern populations. Indeed, the Dorians may have migrated southward to the Peloponnese, across the southern Aegean and Create, and later reached Asia Minor."Thoughts on this? "One of the biggest surprises in the new Lazaridis et al. preprint is that the Natufians don't show any Sub-Saharan African admixture when poked and prodded directly with formal statistics. However, TreeMix, which runs on formal statistics, doesn't have much trouble finding Sub-Saharan or related ancestry in both the Natufians and Neolithic farmers from the Levant. So what's going on?" eurogenes.blogspot.com/2016/06/the-natufian-puzzle.html
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Post by clouddesignc7 on Jan 24, 2017 9:19:19 GMT -5
So on haplogroup J, a haplogroup J summary
So, as far as haplogroup J in general goes (M304, 12f2, etc) in general, so as far as J in general goes, over 90% of J in Egypt / North Africa dates from the Arab expansion into North Africa in the 7th Century C.E. / A.D., and so gets there AFTER or POST Pharaohnic times.
Of J-M267, 90% or more of it's attributed to the 7th Century Arab expansions, and J-M172 is generally seen as being recent admixture.
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Post by clouddesignc7 on Jun 17, 2019 16:46:59 GMT -5
testing...
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Post by clouddesignc7 on Jun 17, 2019 16:47:41 GMT -5
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